Zalmoxes

Dinosaur
Fossil of Zalmoxes in Brussels
Fossil of Zalmoxes in Brussels
Zalmoxes was a small, bipedal ornithopod dinosaur that lived in what is now Romania during the late Cretaceous, around 70 to 66 million years ago, until the asteroid impact that wiped out the non-avian dinosaurs. The name comes from the Dacian deity Zalmoxis.

Where Did Zalmoxes Live?

Zalmoxes inhabited Hațeg Island, a large, isolated landmass in the region, during a time when much of Europe was a series of islands scattered throughout a warm, tropical sea known as the Tethys. Hațeg Island, roughly the size of Ireland, featured woodlands, rivers, open plains, and a warm, subtropical climate. Its long isolation led to the evolution of many unique species.

The island's wildlife included small, distinctive dinosaurs and other unusual creatures. Many dinosaurs - such as the sauropods Magyarosaurus and Paludititan, the duck-billed Telmatosaurus, and Zalmoxes itself - were much smaller than their mainland relatives. Other inhabitants included armored dinosaurs, small carnivorous dinosaurs, crocodiles, tiny mammals, snakes, and lizards.

Species and Size

Size of several dinosaurs compared to a human, with Zalmoxes in purple. Click to enlarge.
Size of several dinosaurs compared to a human, with Zalmoxes in purple. Click to enlarge.
Skull of Zalmoxes
Skull of Zalmoxes
Currently, there are two known species: Zalmoxes robustus and Zalmoxes shqiperorum. The two species looked fairly similar, with Z. shqiperorum being somewhat larger - by up to around 50 cm. However, since the specimens of Z. robustus we have are thought to be sub-adults, it is possible they may have reached larger sizes and closed this gap a little more. Except where otherwise specified, this article will focus mainly on the type species - Z. robustus.

Current specimens range in length from around 2 - 2.4 m, and it is thought that an adult would have reached up to 2.5 m, with a weight of around 45 kg.

What Did Zalmoxes Look Like?

Zalmoxes is known from a skull that is about 80% complete, though most of the skull bones have been found separately rather than joined together. Despite this, paleontologists have reconstructed the general shape of its head, which was deep and triangular, and relatively large for its body size. Zalmoxes also had a stout, robust body with strong limbs and a deep chest, matching the meaning of its species name, "robustus." The backbone is well represented in the fossil record, with bones from the neck, back, and tail, including a sacrum made up of three vertebrae. While many hand and foot bones are missing, the rest of the skeleton shows that Zalmoxes was a small, sturdily built plant-eater.

There is currently no evidence that Zalmoxes had feathers. Zalmoxes belonged to a group of plant-eating dinosaurs called ornithopods, which are part of a larger family known as ornithischians - mostly herbivorous dinosaurs that include famous examples like Stegosaurus and Triceratops. While a few distant ornithischians had simple, hair-like filaments, no such features have ever been found in Zalmoxes or its closest relatives. Zalmoxes likely had scaly skin, much like a traditional reptile.

What Did Zalmoxes Eat?

The teeth of Zalmoxes were typical of its group, the rhabdodontids. It had a beak at the front of the mouth and rows of spade-shaped teeth behind it, well suited for eating tough, fibrous plants like flowering plants, ferns, and horsetails. Studies of its jaw and teeth show that Zalmoxes likely fed by cropping and chewing sturdy vegetation, using its strong jaws and distinctive teeth to process its food.

Discovery and Naming

Zalmoxes was first discovered in Transylvania, Romania, where many fossils were found and originally named Mochlodon robustus in 1899, referring to its sturdy build. Over time, the species was renamed to Rhabdodon robustum, and later research showed it was different enough from Rhabdodon to warrant its own genus. The name Zalmoxes was chosen after a Thracian deity, reflecting the fossil's "resurrection" from the ground into scientific knowledge. A second species, Zalmoxes shqiperorum, was also named at this time in honor of Albania. Some fossils first thought to belong to Zalmoxes have since been reassigned to the related dinosaur Telmatosaurus.

Did Zalmoxes Evolve through Island Dwarfism?

Zalmoxes lived on what was once an island, and for many years scientists believed it was small because of "island dwarfism" - a process where animals living on islands often evolve to become smaller than their relatives on the mainland, usually due to limited space and food. However, more recent research suggests that Zalmoxes may not be a true example of this trend. Instead, its small size appears to match that of its earliest ancestors, while some of its relatives either grew larger or smaller after adapting to different environments. Studies of its bones also show that Zalmoxes grew slowly over a long period, suggesting it may have followed a unique growth strategy rather than simply shrinking in response to island life.

Predators and Defense

Illustration of Hatzegopteryx preying on Zalmoxes
Illustration of Hatzegopteryx preying on Zalmoxes
Given its small size and lack of armor, Zalmoxes would have been vulnerable to a range of predators on Hațeg Island. Fossil evidence shows that threats included large predatory reptiles and snakes, as well as giant pterosaurs, such as the enormous Hatzegopteryx, which is considered the island's top predator and may have preyed on smaller dinosaurs like Zalmoxes.

There is no direct evidence for how Zalmoxes protected itself from predators, so most suggestions are based on comparisons with similar animals. It is possible that Zalmoxes stayed close to dense vegetation for cover or lived in groups for safety, but these behaviors have not been confirmed in the fossil record and remain speculative.

References & Attributions Image: Fossil of Zalmoxes in Brussels - Ghedoghedo, CC BY-SA 3.0, via Wikimedia Commons

Image: Size of several dinosaurs compared to a human, with Zalmoxes in purple. Click to enlarge. - User:Slate Weasel, Public domain, via Wikimedia Commons

Image: Skull of Zalmoxes - Ghedoghedo, CC BY-SA 3.0, via Wikimedia Commons

Image: Illustration of Hatzegopteryx preying on Zalmoxes - Mark Witton, CC BY-SA 4.0, via Wikimedia Commons